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Author notes

*

School of Biosciences, Geoffrey Pope Building, University of Exeter, Stocker Road, Exeter EX4 4QD, UK. E-mail: [email protected].

**

Department of Mathematics and Statistics, University of Lancaster, Lancaster LA1 4YF, UK. E-mail: [email protected].

PHS Consulting Limited, Pryors Hayes Farm, Willington Road, Oscroft, Tarvin, Chester CH3 8NL, UK.

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