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Table 1. 

Summary of inputs and outputs of the control network. Each cell has a copy of the same network, but receives different signals to its inputs.

TypeNameDescription
Input Time Linearly scales from 0 to 1 until the final developmental step is reached. 
X position Horizontal coordinate of physical position of the cell during development. 
Y position Vertical coordinate of physical position of the cell during development. 
Neighbors' morphogen 1 Average value of morphogen output 1 of neighboring cells. 
Neighbors' morphogen 2 Average value of morphogen output 2 of neighboring cells. 
Bias Fixed value 1. 
 
Output Inhibit division Prevents divisions of the cell if above 0. 
Relative division angle Determines the angle at which the cell will divide and its future orientation vector. Output value is multiplied by π and interpreted as a relative angle from −π to π. The final division angle is relative to the mean angle of neighboring cells' orientation. 
Die If above 0, the cell is removed from the embryo. 
Morphogen 1 The state of this output can be indirectly accessed by the cell's neighbors. 
Morphogen 2 Same as above. 
Oscillation period The value of this output at the end of development is used to determine the frequency of contractions of springs attached to the cell. 
Oscillation phase shift The value of this output at the end of development is scaled to the range (−π, π) and determines the phase shift of contractions of springs attached to this cell. 
TypeNameDescription
Input Time Linearly scales from 0 to 1 until the final developmental step is reached. 
X position Horizontal coordinate of physical position of the cell during development. 
Y position Vertical coordinate of physical position of the cell during development. 
Neighbors' morphogen 1 Average value of morphogen output 1 of neighboring cells. 
Neighbors' morphogen 2 Average value of morphogen output 2 of neighboring cells. 
Bias Fixed value 1. 
 
Output Inhibit division Prevents divisions of the cell if above 0. 
Relative division angle Determines the angle at which the cell will divide and its future orientation vector. Output value is multiplied by π and interpreted as a relative angle from −π to π. The final division angle is relative to the mean angle of neighboring cells' orientation. 
Die If above 0, the cell is removed from the embryo. 
Morphogen 1 The state of this output can be indirectly accessed by the cell's neighbors. 
Morphogen 2 Same as above. 
Oscillation period The value of this output at the end of development is used to determine the frequency of contractions of springs attached to the cell. 
Oscillation phase shift The value of this output at the end of development is scaled to the range (−π, π) and determines the phase shift of contractions of springs attached to this cell. 
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