Table 9. 
This table compares the two forms of external fitness, elite seed fitness compared with random seeds of the same size and density (see Figures 6 and 10 in Sections 4.1 and 4.2) versus current winner compared with past winners (see Figure 11).
 Fitness scores averaged over all generations
External fitness based on comparison with past winnersExternal fitness based on comparison with random seeds
Layer 1 2.3 0.739 
Layer 2 3.2 0.814 
Layer 3 2.4 0.799 
Layer 4 11.7 0.892 
Layer 4 Shuffled 10.8 0.871 
Layer 4 Mutualism 12.0 0.897 
Correlation 0.767 
p-value of correlation 4.15E−15 
p-value < 0.05 Yes 
 Fitness scores averaged over all generations
External fitness based on comparison with past winnersExternal fitness based on comparison with random seeds
Layer 1 2.3 0.739 
Layer 2 3.2 0.814 
Layer 3 2.4 0.799 
Layer 4 11.7 0.892 
Layer 4 Shuffled 10.8 0.871 
Layer 4 Mutualism 12.0 0.897 
Correlation 0.767 
p-value of correlation 4.15E−15 
p-value < 0.05 Yes 

Notes. The table shows the correlation between the two measures of external fitness. The correlation is calculated from 72 random seed fitness scores and 72 past winner fitness scores (6 layers with 12 fitness values for each layer yields 72 fitness scores). Each fitness score in this table is an average over all the generations for a given layer (an average of 12 fitness values, one for each run, each of which is an average over the 100 generations). We evaluate the statistical significance of the correlations using a two-tailed Student t-test for Pearson correlations. The correlation is high (0.767), and it is statistically significant.

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