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Table 10.

Like Table 9, this table compares the two forms of external fitness: elite seed fitness compared with random seeds of the same size and density, versus current winner compared with past winners.

	Fitness scores in generation 100
	External fitness based on comparison with past winners	External fitness based on comparison with random seeds
Layer 1	2.7	0.743
Layer 2	7.7	0.846
Layer 3	4.4	0.810
Layer 4	21.6	0.936
Layer 4 Shuffled	24.5	0.927
Layer 4 Mutualism	23.6	0.944
Correlation	0.765
p-value of correlation	5.37E−15
p-value < 0.05	Yes

	Fitness scores in generation 100
	External fitness based on comparison with past winners	External fitness based on comparison with random seeds
Layer 1	2.7	0.743
Layer 2	7.7	0.846
Layer 3	4.4	0.810
Layer 4	21.6	0.936
Layer 4 Shuffled	24.5	0.927
Layer 4 Mutualism	23.6	0.944
Correlation	0.765
p-value of correlation	5.37E−15
p-value < 0.05	Yes

Notes. Each fitness score in this table is an average of the final generation for a given layer (an average of 12 fitness values). The difference between the two tables is that here we focus on the final generation, whereas the preceding table examined the average fitness over all the generations. The similar correlation values in Table 9 (0.767) and Table 10 (0.765) indicate that the correlations between the two forms of external fitness (comparison with past winners and comparison with random seeds) are robust.

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